top of page


You can find our publications on Google Scholar

Xia, Y, J. Sanderman, J.D. Watts, M.B. Machmuller, A.L. Mullen, C. Rivard, A. Endsley, H. Hernandez,

J. Kimball, S.A. Ewing, M. Litvak, T. Duman, P. Krishnan, T. Meyers, N.A. Brunsell, B. Mohanty, H. Liu, Z. Gao, J. Chen, M. Abraha, R.L. Scott, G.N. Flerchinger, P.E. Clark, P.C. Stoy, A.M. Khan, E.N. J. Brookshire, Q. Zhang, D.R. Cook, T. Thienelt, B. Mitra, M. Mauritz-Tozer, C.E. Tweedie, M.S. Torn, D. Billesbach. 2024. Coupling remote sensing with a process model for the simulation of rangeland carbon dynamics. Under review.

Stewart, R.D., M. Flury, H. Ajami, R.G. Anderson, T.R. Green, Y. Jin, A. Patrignani, R. Shillito, W. Zhang,

M.R.A. Najm, E. Babaeian, M. Berli, E. N. J. Brookshire, A.L.M. Daigh, J. Giovando, R. Heinse, J. Heitman, J. Huang, T. Kelleners, M. Naseri, T.E. Ochsner, J. Radolinski, M. Sadeghi, S. Sasidharan, M.K. Shukla, M.Tuller, O. Wendroth, J.Wu, B.M. Wyatt, Y. Yang, Y. Yu, F. Zhang. 2024. Emerging issues and research opportunities in vadose zone processes. Under review.


Gay, J.D*., B. Currey*, K.T. Davis, E.N.J. Brookshire. 2024. Functional attributes of conifers expanding

into semi-arid grasslands modulate carbon and nitrogen fluxes in response to prescribed fire.  Submitted to Biogeochemistry.

Hu, C-C, A.W. Driscoll, Y-W. Kuang, E. N. J. Brookshire, X-T. Lü, C-J. Chen, W. Song1, C-Q. Liu, B.Z.

Houlton, X-Y Liu. 2024. Surface temperature regulated global nitrogen-use patterns of terrestrial plants. Under review.


Abeshu, G.W*. , H.Y. Li , M. Shi, E.N.J. Brookshire, J. Tang, C. Xu, N. McDowell, L.R. Leung. 2024. A

generalized relationship linking water balance and vegetation carbon uptake across site-to-regional scales. Journal of Hydrologic Engineering.  


Wendt, J.A.F.*, D.B. McWethy, G.P. Hempson, E.N.J. Brookshire, S.D. Fuhlendorf. 2023. Past and present

biomass consumption by herbivores and fire across productivity gradients in North America. Environmental Research Letters.


Pellegrini, A.F.A., P.B. Reich, S.E. Hobbie, C. Coetsee, B. Wigley, E. February, K. Georgiou, C. Terrer,

E. N. J. Brookshire, A. Ahlström, L. Nieradzik, S. Sitch, J. Melton, M. Forrest, F. Li, S. Hantsen, C. Burton, C. Yue, P. Ciais, R.B. Jackson. 2023. Soil carbon storage capacity of drylands under altered fire regimes. Nature Climate Change.


Poulter, B., B. Currey*, L. Calle, A.N. Shiklomanov, C.H. Amaral, E.N.J. Brookshire, P. Campbell , A.

Chlus, K. Cawse‐Nicholson, F. Huemmrich, C. E Miller, K. Miner , Z. Pierrat, A.M. Raiho, D. Schimel, S. Serbin, N. Stavros, J. Stutz, P. Townsend, D.R. Thompson, Z. Zhang. 2023. Simulating global dynamic surface reflectances for imaging spectroscopy spaceborne missions ‐ LPJ‐PROSAIL. JGR-Biogeosciences.


Gay, J.D.*, Currey, B.*, E.N.J. Brookshire. 2022. Global distribution and climate sensitivity of the tropical

 montane forest N cycle. Nature Communications.


Gay, J.D.*, H.M. Goemann*, B. Currey*, P.C. Stoy, J.R. Christiansen, P.R. Miller, B. Poulter, B.M. Peyton,

E.N.J. Brookshire. 2022. Climate mitigation potential and soil microbial response of cyanobacteria-fertilized bioenergy crops in a cool semi-arid cropland. GCB-Bioenergy.


Stoy, P.C., K. Van Dorsten, A. Khan, P. Sauer, T. Weaver, E.N.J. Brookshire. 2022. Ecosystem gross primary

productivity after autumn snowfall and melt events in a mountain meadow. JGR-Biogeosciences.


Currey, B.*, D.B. McWethy, N. Fox, E.N.J. Brookshire. 2022. Large contribution of woody plant expansion

to recent vegetative greening of the Northern Great Plains. Journal of Biogeography. doi: 10.1111/jbi.14391


Sigler, W.A.*, S.A. Ewing, S.D. Wankel, C.A. Jones, S. Leuthold, E.N.J. Brookshire, R.A. Payn. 2022.

Isotopic signals in an agricultural watershed suggest denitrification is locally intensive in riparian areas but extensive in upland soils. Biogeochemistry.


Rodríquez-Cardona, B., A.S. Wymore, A. Argerich, R.T. Barnes, S. Bernal, E.N.J. Brookshire, A. A. Coble,

W.K. Dodds, H.M. Fazekas, A.M. Helton, P.J. Johnes, S.L. Johnson, J.B. Jones, S.S. Kaushal, P. Kortelainen, C. Lopez-Lloreda, R.G. M. Spencer, W.H. McDowell. 2022. Shifting stoichiometry: Long-term trends in stream dissolved organic matter reveal altered C:N ratios due to history of atmospheric acid deposition. Global Change Biology. doi: 10.1111/GCB.15965


Goemann, H.M.*, J.D. Gay*, B.M. Peyton, R.C. Mueller, E.N.J. Brookshire, B. Peyton, B. Poulter. 2021.

Above- and below ground responses to field applications of cyanobacterial biofertilizer in a semi-arid bioenergy cropping system. GCB-Bioenergy. doi: 10.1111/gcbb.12892


Wymore, A.S., P.J. Johnes, S. Bernal, E.N.J. Brookshire, H.M. Fazekas, A.M. Helton, A. Argerich, R.T.

Barnes, A. A. Coble, W.K. Dodds, S. Haq, S.L. Johnson, J.B. Jones, S.S. Kaushal, P. Kortelainen, C. Lopez-Lloreda, B. Rodríquez-Cardona, R.G. M. Spencer, P.L. Sullivan, C. A. Yates, W.H. McDowell. 2021. Gradients of anthropogenic nutrient enrichment alter N composition and DOM stoichiometry in freshwater ecosystems. Global Biogeochemical Cycles.


Epstein, K., J. Haggerty, K. Roemer, B. Currey*, H. Duff, J. Gay*, H. Goemann*, S. Loewen, M. Milligan, J.   Wendt*, D. Wood, E.N.J. Brookshire, L. McNew, D. McWethy, P. Stoy, B. Maxwell. 2021. Towards an

urgent yet deliberate conservation strategy: sustaining social-ecological systems in rangelands of the Northern Great Plains, Montana. Ecology and Society.


Currey, B.*, M.P. Oatham, E.N.J. Brookshire. 2021. Negative trait-based association between tropical forest

biomass accumulation and abundance of nitrogen fixing trees. Journal of Ecology.


Stoy, P.C., A.A. Cook*, J.E. Dore, W. Kleindl, E.N.J. Brookshire, T. Gerken. 2021. Methane efflux from an

American bison herd. Biogeosciences,

Brookshire, E. N. J., P. C. Stoy, B. Currey, and B. Finney. 2020. The greening of the Northern Great Plains and its biogeochemical precursors. Global Change Biology 26:5404–5413.

Valiela, I., R. Juman, H. Asmath, D. Hanacek, J. Lloret, E. Elmstrom, K. Chenoweth, and E. N. J. Brookshire. 2020. Water quality, nutrients, and stable isotopic signatures of particulates and vegetation in a mangrove ecosystem exposed to past anthropogenic perturbations. Regional Studies in Marine Science 35:101208.

Brookshire, E. N. J., N. Wurzburger, B. Currey, D. N. L. Menge, M. P. Oatham, and C. Roberts. 2019. Symbiotic N fixation is sufficient to support net aboveground biomass accumulation in a humid tropical forest. Scientific Reports 9:7571.

Sigler, W.A, S.A. Ewing, C.A. Jones, R.A. Payn, E.N.J. Brookshire, J.K. Klassen, D. Jackson-Smith, G.S. Weissmann. 2018. Connections among soil, ground, and surface water chemistries characterize nitrogen loss from an agricultural landscape in the upper Missouri River basin.  Journal of Hydrology 556: 247-261

Stoy, P.C., S. Ahmed, M. Jarchow, B. Rashford, D. Swanson, S. Albeke, G. Bromley, E.N.J. Brookshire, M. Dixon, J. Haggerty, J. Kerby, P. Miller, B. Peyton, T. Sohl, L. Spangler, C. Straub, A. Royem, B Poulter. 2018. Opportunities and tradeoffs between bioenergy with carbon capture and storage (BECCS) and food, water, energy, biodiversity, and social systems at regional scales. BioScience 68: 100-111

Wurzburger, N., and E. N. J. Brookshire. 2017. Experimental evidence that mycorrhizal nitrogen strategies affect soil carbon. Ecology 98:1491–1497.

Brookshire, E. N. J., S. Gerber, W. Greene, R. T. Jones, and S. A. Thomas. 2017. Global bounds on nitrogen gas emissions from humid tropical forests. Geophysical Research Letters 44:2502–2510.

Lajtha, K, Bai, E, Baisden, T, Berhe, AA, Bowden, B, Brookshire, E.N.J., Brozostek, E, Crow, S, Driscol, C, Evans, C, Finlay, J, Fisk, M, Grandy, S, Hamdan, LJ, Harrison, J, Hawkes, C, Kalbitz, K, Kaushal, S, Kramer, M, Matzner, E, Melack, J, Mulder, J, Porder, S, Sanderman, J, Stanley, E, Tank, J, Vile, M, Voss, M, Wieder, K, Ziegler. 2017. Brave new world: Editorial in support of environmental science. Biogeochemistry. 133:3–5

Wurzburger, N., E. N. J. Brookshire, M. L. McCormack, and R. A. Lankau. 2017. Mycorrhizal fungi as drivers and modulators of terrestrial ecosystem processes. New Phytologist 213:996–999.

Gücker, B., R. C. S. Silva, D. Graeber, J. A. F. Monteiro, E. N. J. Brookshire, R. C. Chaves, and I. G. Boëchat. 2016. Dissolved nutrient exports from natural and human-impacted Neotropical catchments. Global Ecology and Biogeography 25:378–390.

Craine, J. M., E. N. J. Brookshire, M. D. Cramer, N. J. Hasselquist, K. Koba, E. Marin-Spiotta, and L. Wang. 2015a. Ecological interpretations of nitrogen isotope ratios of terrestrial plants and soils. Plant and Soil 396:1–26.

Brookshire, E. N. J., and T. Weaver. 2015. Long-term decline in grassland productivity driven by increasing dryness. Nature Communications 6:7148.

Yano, Y., E. N. J. Brookshire, J. Holsinger, and T. Weaver. 2015. Long-term snowpack manipulation promotes large loss of bioavailable nitrogen and phosphorus in a subalpine grassland. Biogeochemistry 124:319–333.

Craine, J. M., A. J. Elmore, L. Wang, L. Augusto, W. T. Baisden, E. N. J. Brookshire, M. D. Cramer, N. J. Hasselquist, E. A. Hobbie, A. Kahmen, K. Koba, J. M. Kranabetter, M. C. Mack, E. Marin-Spiotta, J. R. Mayor, K. K. McLauchlan, A. Michelsen, G. B. Nardoto, R. S. Oliveira, S. S. Perakis, P. L. Peri, C. A. Quesada, A. Richter, L. A. Schipper, B. A. Stevenson, B. L. Turner, R. A. G. Viani, W. Wanek, and B. Zeller. 2015b. Convergence of soil nitrogen isotopes across global climate gradients. Scientific Reports 5:8280.

Thomas, R. Q., E. N. J. Brookshire, and S. Gerber. 2015. Nitrogen limitation on land: how can it occur in Earth system models? Global Change Biology 21:1777–1793.

Gerber, S. and E.N.J. Brookshire.  2014.  Scaling of physical constraints at the root-soil interface to macroscopic patterns of nutrient retention in ecosystems.  American Naturalist 183:418-430.

Brookshire, E. N. J., and S. A. Thomas. 2013. Ecosystem Consequences of Tree Monodominance for Nitrogen Cycling in Lowland Tropical Forest. PLoS ONE 8.

Brookshire, E. N. J., L. O. Hedin, J. D. Newbold, D. M. Sigman, and J. K. Jackson. 2012b. Sustained losses of bioavailable nitrogen from montane tropical forests. Nature Geoscience 5:123–126.

Brookshire, E.N.J., S. Gerber, D.N.L. Menge, and L.O. Hedin.  2012. Large losses of inorganic nitrogen from tropical rainforests suggest a lack of nitrogen limitation.  Ecology letters 15: 9-16.

Brookshire, E.N.J., S. Gerber, J.R. Webster, J. Vose, and W. Swank. 2011.  Direct effects of temperature on forest nitrogen cycling revealed through analysis of long-term watershed records. Global Change Biology 17:297-308.

Hedin, L.O., E.N.J. Brookshire, D.N.L. Menge, and A.R. Barron. 2009. The nitrogen paradox in tropical forest ecosystems. Annual Review of Ecology, Evolution, and Systematics 40: 613-636.

Brookshire, E.N.J., H.M. Valett, and S. Gerber. 2009. Maintenance of terrestrial nutrient loss signatures during in-stream transport. Ecology 90:293-299.

Brookshire, E.N.J., H.M. Valett, S.A. Thomas, and J.R. Webster. 2007. Atmospheric N deposition increases organic N losses from temperate forests. Ecosystems 10:252-262.

Brookshire, E.N.J., H.M. Valett, S.A. Thomas, and J.R. Webster. 2005. Coupled cycling of dissolved organic nitrogen and carbon in a forest stream. Ecology 86: 2487-2496.

Kauffman, J.B., A.S. Thorpe, and E.N.J. Brookshire. 2004.  Livestock exclusion and belowground ecosystem responses in riparian meadows of eastern Oregon.  Ecological Applications 14: 1671-1679.

Dwire, K.A., J.B. Kauffman, E.N.J. Brookshire, and J. Baham. 2004.  Plant biomass and species composition along an environmental gradient in montane riparian meadows.  Oecologia 139: 309-317.

Brookshire, E.N.J., and K.A. Dwire. 2003.  Controls on patterns of coarse organic particle retention in headwater streams.  Journal of the North American Benthological Society 22:17-34.

Brookshire, E.N.J., J.B. Kauffman, D. Lytjen, and N. Otting. 2002. Cumulative effects of wild ungulate and livestock herbivory on riparian willows. Oecologia 132:559-566.

bottom of page